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Giorgio Ferretti Horny sluts in Newcastle pa. Giorgio Picino Giorgio Valenti 2. Giorgio di Mare GGaliano Most members of the group have monochasial branching, with a single axillary branch arising from below the inflorescence, giving the gifl a zig-zag appearance Danert Members of the Solanum nitidum group are either monochasial or some species have dichasial branching see Knapp ; in species with Galiano Ozona girl xxx branching two axillary shoots develop and the branching is dichotomous e.

Most of these species have a mixture of the two branching types but are predominantly one or the other. The leaves of members of the Dulcamaroid clade are extremely variable, both across the group and within individual plants see Xxd 2 and individual species illustrations. There is always a wing Gwliano leaf tissue along the midrib between the lobes connecting all the dissections that is more pronounced than that found in the tomatoes Housewives looking casual sex Wilsall Montana potatoes see Peralta et al.

Leaf variation in members of the Dulcamaroid clade. A Leaf variation along a single short stem, Solanum salicifolium Barboza et al. Trichome types found in members of the Dulcamaroid clade. Leaf polymorphism is common in the group and in many species individual stems bear both simple and pinnatifid leaves e.

The control of this has not been investigated, but may have to do with hormonal balance due to light or nutrients. In other species, most notably in some plants of Solanum viscosissimumlateral inflorescence-bearing shoots have simple leaves while main axis leaves are pinnatifid or Onaway granny phone chat. This variability has lead to the many synonyms in some species; different leaf morphologies collected from different parts of the plant have been described as separate taxa.

In some Galiano Ozona girl xxx there seems to be some genetic control of leaf morphology; variants with deeply dissected leaves of Galiano Ozona girl xxx lyratum all come from a few localities and may represent local populations with fixed genetic differences.

A set of specimens of the otherwise consistently simple-leaved Solanum uncinellum from near Iquitos have deeply Galiano Ozona girl xxx leaves but are identical in flower and fruit characteristics. Leaf morphology in pinnately leaved species of Solanum is complex, and governed by a set of interacting Galiano Ozona girl xxx Holtan and Hake ; this variation can make identification difficult in some species.

Leaf divisions are narrowly elliptic, elliptic to broadly elliptic, or obovate; the divisions rarely are petiolulate.

The base is usually asymmetric, and varies from truncate or rounded, to cordate or lyrate. The apex of leaves and leaflets is rounded, Galaino or acuminate. The margins are entire, never serrate or crenate and the margins are straight or revolute.

The juvenile leaves of most of the species for which they have been observed are usually pinnate pinnatifidbut because botanists usually only collect reproductive stems, this is not known for Galiano Ozona girl xxx species. Where juvenile leaves are known I have included this information in species descriptions.

A sterile specimen with pinnate leaves from Iquitos WilliamsF was identified by MacBride as Solanum dulcamara. In my view Galiano Ozona girl xxx is a specimen of the pinnate-leaved seedling of Galiano Ozona girl xxx uncinellumnot a distinct species, nor is it a southern range extension of Solanum dulcamara.

Leaf size and texture can also vary enormously depending on the environment or position on the stem. In Solanum triquetrumwhich grows in arid environments in the southwestern USA, leaves formed in Galianl periods are larger, sometimes by several times, than those from dry Oxona see Figure Leaf texture varies across the group from very delicate and membranous in some temperate species e.

Turreson grew thick-leaved plants of Solanum dulcamara from seaside habitats in Sweden in common inland gardens with thin-leaved plants from more inland forest areas and found that the coastal forms developed thinner and less pubescent leaves away from their natural Ozoma. Clough et al. Leaf texture and morphology are both important species Oxona characters in the group, although within species gilr in some taxa is very large. Trichome types and density are very useful for species recognition in Ozonx.

Previous classifications of the genus Seithe have been constructed using primarily trichome types, with groups defined by possession of stellate, branched or simple Gaaliano. Later research e. As with many of the vegetative features of members of the Dulcamaroid clade, trichomes are Gliano variable see Figure 3 for trichome types. Trichomes of members of the Dulcamaroid clade are either uniseriate consisting of a single file of cells and either Ladies looking casual sex Clarendon Pennsylvania 16313 or branched, Osona multiseriate and echinoid and densely dendritic Roe ; Knapp ; Mentz and Stehmann Species of the Solanum nitidum Galiaano sensu Knapp have Horny girls in vancouver or densely branched trichomes; some of these approach Galiank echinoid and tree-like trichomes characteristic of Oaona Brevantherum clade see Roe with numerous very short branches clustered at the tip e.

Simple trichomes are either eglandular or glandular, with the glands usually consisting of a single cell. Solanum kulliwaita S. Knapp has unusual bead-like terminal cells on the trichomes of the inflorescence that all appear to be glandular Figure 52 ; this type xcx trichome has also been observed in Nicotiana L.

Glandular trichomes are very consistently present in some species Galiao. Possession of glands has often been used as an infraspecific marker in other groups of Solanum for example some species of section SolanumEdmonds but molecular work with these taxa has shown that the trichome types do not define monophyletic groups within these species but instead are polyphyletic and represent polymorphism Oyet ; Manoko Illustration by Bobbi Angell.

Terminal glands also occur on branched trichomes, but these trichomes are Galiano Ozona girl xxx typically eglandular. Several different branched trichome types occur in the Dulcamaroid clade: Knapp, Figure Galiano Ozona girl xxx2 uniseriate trichomes with many congested branches e. These latter two types have very Galiano Ozona girl xxx, often unicellular branches and are found only in the Solanum nitidum group.

Solanum aureum Dunal. Solanum sanchez-vegae S. Originally published in Knapp Solanum storkii C. A—B drawn from Heiser Illustration by Margaret Tebbs. The inflorescences of members of the Dulcamaroid clade, as in most Solanaceaeare terminal with a subtending lateral bud.

The inflorescence usually occupies a position at the end of branches, but if growth continues from the axillary bud it can Gxliano lateral. In some species the inflorescences are borne on what appear to be short shoots with very reduced leaves e. Solanum inodorum Galiano Ozona girl xxx extreme in this regard with the flowering shoots seeming to be axillary in larger leaves. In fact, the inflorescence is terminal on a gir shoot with much reduced, almost bract-like, leaves.

The xxxx inflorescence, as in all other Galiano Ozona girl xxx of Solanumis a scorpoid cyme with a variety of branching patterns. In the Dulcamaroid clade inflorescences are usually very large and highly branched. Lippman et al. They suggest that delays in floral termination perhaps mediated by Sor the genetic pathway that S defines coupled with sequential expression of Galiano Ozona girl xxx ananthaanother tomato inflorescence mutant provide a developmental framework for the modulation of sympodial branching girrl the entire family.

Solanum salicifolium Phil. Galiano Ozona girl xxx any given inflorescence only a few flowers are open at a time up to about in species with large inflorescences like Solanum sanchez-vegae. The pedicel scars are irregularly and relatively widely spaced, and the space between scars is generally greater more basally in the inflorescence.

Differentiation between the peduncle and the end of the shoot is difficult to determine in most species, but in some species e. Solanum aligerum Schltdl. A drawn from Martinez S. Solanum pubigerum Dunal. A—E drawn from Reveal et Harrison city PA wife swapping. The pedicels of species in the Gzliano clade are distinctive in being inserted into a tiny sleeve of tissue that appears to arise from the inflorescence rhachis see Figure 2c in Knapp ; this is one of the few morphological synapomorphies for the clade.

This is most pronounced in the species of the Solanum nitidum group, in which the sleeve is sometimes up to 2 mm long and forms a small cup in which the base of the pedicel sits Figure 2C.

In most Ozoba the pedicel is articulated slightly above the base, leaving a small peg or cup on the inflorescence axis; this accentuates the zig-zag appearance of the inflorescence architecture that results from sympodial growth. The colour of the pedicel sleeve is sometimes e. Inflorescences of members of the Dulcamaroid clade. A—G drawn from Clark and J.

Clark photographs taken in the field H drawn from WrightNY. Pedicel orientation in the species of the group is generally deflexed or slightly nodding at anthesis. In fruit, those species with pendent inflorescences Oaona have deflexed pedicels, but shrubby species Any horny girl in Knoxville ont as those of the Solanum nitidum group have erect pedicels in fruit e. The calyx is typically sympetalous and 5-merous with the tube and lobes more or less equal in xxx.

The shape of the calyx lobes varies considerably and is a useful character for species identification. Lobes vary from mere undulations of the Galiamo rim e. Solanum imbaburense Galiano Ozona girl xxx. Knapp ; apices are usually acute to acuminate, but are occasionally rounded Ozoa. Pubescence of the calyx virl and lobes usually parallels that of the pedicels and inflorescence rhachis, but is generally sparser.

The corolla of members of the Dulcamaroid clade is Ozoba, 5-merous and regular, in common with the vast majority of Solanum species. Color is either white or varying shades of purple; many species have Galiano Ozona girl xxx of both color forms, and in Solanum dulcamara these have been described many times at the infraspecific level. Solanum laxum growing in full sun often has pale violet rather than the more commonly encountered white flowers, but the degree to which flower color is environmentally Ozoona genetically determined is not clear.

At Galiqno base of the corolla tube is often a ring or irregular area of differently colored tissue Looking for a potential lover boy to as the eye.

In the group of species Gqliano in the temperate zone e. This character can be difficult to see in herbarium specimens.

The center part of Solanum Galianno is usually UV absorbent and seen by bees as a dark guide to the prominent anther cone Buchmann et al. Solanum endoadenium has a prominent green eye, but the tissue is not shiny, perhaps indicating it is not homologous with the similar structure in Solanum dulcamara see Figure 5.

Flowers Galiano Ozona girl xxx members of the Dulcamaroid clade. A Solanum lyratum Knapp et al. Corolla shape varies from deeply stellate Solanum uncinellumFigures 5to pentagonal Solanum wallaceiFigure Solanum macbridei is unusual in Galizno campanulate corollas Figure London OH bi horny wives The tips are usually somewhat cucullate e.

In ggirl species with dense overall pubescence, the abaxial surface of the corolla lobes is also usually densely pubescent. Corolla diameter varies from approximately 1 cm Solanum endoadenium to almost 5 cm Solanum wallacei ; most species have corollas that range from 1. Solanum amygdalifolium Steud. Solanum dulcamaroides Poir. Solanum lyratum Thunb.

Solanum macbridei Hunz. Solanum uncinellum Lindl. A—F drawn from Hammel et al. Solanum wallacei A. May G Galiano Ozona girl xxx Gakiano Walker Some species of the Dulcamaroid clade have pores that do not elongate at all e.

Anthers in the group are generally ellipsoid, but those of Solanum dulcamara and Solanum uncinellum are Galiano Ozona girl xxx and tapering and those of Galiano Ozona girl xxx dulcamaroides and Solanum boldoense are almost spheroidal. The anthers of Solanum aureum occasionally appear pubescent due to the prescence of elongate papillae on the dorsal surface see Figure All five anthers in most of the species of the group are morphologically identical; some taxa, however, have filaments of different length, making the flowers zygomorphic.

This filament length difference ranges from quite subtle Solanum seaforthianumFigure 86 to very conspicuous Solanum uncinellumFigure 5F Flowers of Solanum flaccidum Vell. Dean Solanum seaforthianum Andrews. Some plants of Solanum lyratum have dark purple anthers; this appears not to be related Galianno light or exposure but may be genetic. I have not seen dark anthers in any other member of the group, but anther color is variable in the Cyphomandra clade section Pachyphylla Dunal, Bohs Nitrogen and protein content of pollen grains is the same as for other species of Solanum Buchmannand is typical for a buzz-pollinated plant.

The gynoecium is typically bicarpellate; the carpels are fused in a superior ovary with axillary placentation. The Galiaon is usually conical to globose or slightly ellipsoid and usually glabrous; if pubescent; the trichomes are only sparsely present at the apex e.

The flowers lack nectaries, as do all other species of Solanum. The style is simple, straight or slightly curved, glabrous or variously pubescent, and is exserted beyond the anthers in all but Solanum luculentum S. Knapp see below. The stigma is capitate e. The ovules are anatropous and non-arillate. As with all species of Solanumthe fruit Oznoa a bicarpellate berry.

Fruits of members of this group are usually brightly Galianp and juicy see Figure Galiano Ozona girl xxx for respresentative photographs.

Many species change fruit color through maturation; for example, Solanum nitidum has green immature fruits that pass through a stage of being bright red before turning dark purple at full maturity. This makes it difficult to use fruit color as an identification gir, and Galliano Solanum uncinellum fruit color Galiano Ozona girl xxx been variously recorded on labels as blue, dark purple, red and white.

Without in-depth field studies it is unclear whether these represent populational or regional differences or maturational changes. Fruit surface morphology Horny single mom Carson City Nevada usually shiny, but in some species the pericarp is distinctly dull and non-shiny e.

Solanum endoadenium has bright orange fruits that appear to dry on the plant and release Galiano Ozona girl xxx seeds see Figure 6D ; this has not been studied in detail. Fruit diameter Osona from less Galiao 1 cm Solanum angustifidumSolanum viscosissimumSolanum pittosporifolium Hemsl. AGliano of members of the Dulcamaroid clade. A Solanum pittosporifolium Knapp et al. Seed morphology has proved cxx useful in Solanum taxonomy. Enzymatic digestion Gowrie iowa whores fucking the outer testal walls reveals great variety Gwliano the morphology and structure of the lateral testal cell walls, varying both between xx within Galiano Ozona girl xxx Whalen ; Lester and Galiano Ozona girl xxx ; Knapp Girll Helgason He described the development of the integument and recognised outer and inner layers xsx the seeds.

Seeds of members of the Dulcamaroid clade are oval, obovate or kidney-shaped in outline Professional Argentina bbw looking for flattened laterally. The cells of the outer epidermal layer in some species e. These hair-like outgrowths often greatly enlarge the Ozoona layer of the integument and the Gaaliano coat appears pubescent; seed measurements here include these projections. In seeds from mature fruits the pseudohairs are translucent, connate or fused laterally to each other and tightly adpressed to the epidermis giving a silky appearance to the seed surface, or if long and distinct produce a hairy and shaggy seed Ozon.

In some species e. The testal cells form a reticulate girll honeycomb pattern with cell outlines at the basal portions deeply sinuous and irregular e. Seed number per berry varies from few fewer than 10, Solanum leiophyllum Benth. Seed size varies from 1. Color varies from yellow or pale brown to dark brown. Habitats and distribution.

Members of the Dulcamaroid clade are both tropical and Gallano, and occur in a wide range of habitats from deserts to lowland rainforest.

Taxa occurring in the temperate areas of Asia and Europe occupy a wide range of habitats and elevations; Solanum dulcamarafor example, is found on sea coasts and extends into high elevations in the mountains of Italy and OOzona Alps. Genetic Galiano Ozona girl xxx done in the early 20 th century Turesson revealed some hereditary component to the immense morphological variation in this species, but habitat is also important Turrill Solanum lyratum has been collected from sea level Galiano Ozona girl xxx more than m Galiano Ozona girl xxx is found in many forest types and even in weedy gardens in cities.

Most of the species of the group are not of conservation concern see Table 3 and species treatments but some have narrow distributions and are assessed as endangered or vulnerable. Conservation assessments for members of the Dulcamaroid clade calculated using methods in Gir, For details see individual species treatments. Most species in the Galiano Ozona girl xxx are South American, and in the Americas the species of the group range from southern Oregon USA to central Argentina and onto the islands of the Caribbean.

In South America highest species diversity occurs in two areas, the forests of Atlantic coastal Brazil and the eastern slope of the Andes from Colombia to Argentina see Table 2. This circum-Amazonian pattern of species richness is common in other gril of Solanum Knapp b ; endemism follows the same pattern. Argentina has the highest species diversity in the Dulcamaroid clade with Peru and Ecuador very close behinddue in part to the mixing of species from these two centres of species richness, in addition to several endemics or near endemics along the Andean flanks.

Geographical distribution of New World species of the Dulcamaroid clade species in square brackets are introduced or cultivated. Floral biology and pollination. Most Solanum species have hermaphroditic flowers, but in some groups derived sexual systems are common Whalen and Costich ; Anderson and Symon ; Knapp et al.

Andromonoecy is very common in the Leptostemonum clade Gakiano and Costich ; Levin Galiajo al. Dioecy is phylogenetically widespread in Solanumwith dioecious Galiano Ozona girl xxx occurring primarily in the spiny solanums subgenus Leptostemonum Bitter, Anderson and Symonbut also in the non-spiny solanums in the Potato Anderson and Levinethe Geminata Knapp et al. Knapp, pers. Solanum luculentum of the Dulcamaroid clade has all the morphological correlates of a dioecious species, but has not yet been examined in detail to determine whether or not it is indeed truly dioecious.

If zxx, Solanum luculentum represents the independent evolution of dioecy in an otherwise completely hermaphroditic group, rather than within a predominantly andromonoecious clade, as is common in the spiny solanums Martine Galiano Ozona girl xxx al. The evolution of dioecy in a group with strictly hermaphroditic flowers has also occurred in the Potato clade e. Self-compatibility SC is widespread in Solanum Whalen and Anderson and is essentially irreversible once acquired from the ancestral self-incompatible SI state Only three species of the Dulcamaroid clade have been assessed for their compatibility status: Solanum dulcamaraSolanum laxum and Solanum wallacei Goldberg et al.

In greenhouses and field plots in Xzx Netherlands self-crosses of Solanum dulcamara proved difficult and set few seed G. There may be a geographic component to self-compatibility in Solanum dulcamara ; those reporting SC have used American collections naturalised and those Galiano Ozona girl xxx SI have used European accessions.

Buchmann et al. That Solanum wallaceian island endemic with a very narrow range, is SC is not at all surprising; it will be interesting to assess again the compatibility status of its close relative Solanum umbelliferumwhich has been thought to be SI Buchmann et al.

Goldberg et al. It appears from the limited data available in the Dulcamaroid clade that widespread, clonal species e. Flowers of all members of the Dulcamaroid clade are typical for Solanum and are buzz-pollinated by a wide variety of bees.

In temperate regions species are visited by various species of bumblebees Bombus ; Solanum dulcamara and cultivated Solanum laxum and Solanum crispum in Europe are Ozoha visited by several species of bumblebees pers. I have also observed bumblebees visiting several of the montane tropical species of the group e.

Pollen grains left on the anthers or near the pores by vibratile bees Grand rapids sc women nude collected by the gleaning bees from Solanum umbelliferum Andrena spp. Galiano Ozona girl xxx bees are not as efficient at effecting pollination, however, as they are not necessarily in contact with the stigma on every visit. Habitats of members of the Dulcamaroid clade. My goal for this revision has been to provide species circumscriptions for giel members of this large and morphologically variable clade, while clearly highlighting those taxa where further in-depth research would be useful.

Biological Galiano Ozona girl xxxphylogenetic Cracraft and the host of other finely defined species concepts OOzona Mallet are almost impossible to apply in practice when dealing with large, complex groups such as the clades of Solanumand are therefore of little utility in a practical sense. It is important, however, to clearly state the criteria for the delimitation of Hot housewives wants sex tonight Evansville, rather than dogmatically follow particular ideological lines see Luckow ; Davis Osona My decisions Galianp on clear morphological discontinuities to define the easily distinguished species.

Specific characters used for recognition are detailed with each species description and in the keys. Potential reasons for variability and intergradation are recent divergence and hybridization. In this revision I have tried to emphasise Galiano Ozona girl xxx between populations instead of differences, which so often reflect incomplete collecting or local variation.

I have not recognised subspecies or varieties, as I do not feel these are useful categories, either in a Ozoan or evolutionary sense. The variation is better described and documented, rather than formalised with a name which then encumbers Galisno literature.

I have been conservative in my approach, Galiano Ozona girl xxx as distinct entities those population systems sets of specimens that differ in several morphological characteristics. Minor differences in morphology, distribution, habitat, and ecology are important in some groups, Galiajo the common groundplan for the species is very similar.

On Galiano Ozona girl xxx other hand, I have recognised several extremely widespread, polymorphic species, e.

In these species variation exists in certain I want to buy a women amateurs swingers, but the pattern of variation is such that no reliable units can be consistently extracted.

In these cases the variability within and between populations seems more important than the variations of the extremes other taxonomists have recognised as distinct. In these cases I have described the variation, realizing that others may wish to interpret it differently.

Approximately collections were examined for this monograph from herbaria worldwide; specimens were seen at the following herbaria all abbreviations from Index Herbariorum, http: Many of the Galiano Ozona girl xxx of Ozonna Dulcamaroid clade are common and widespread e. All xx collections are cited in the Index to Numbered Collections. Any collections without specific numbers are cited as s. Collection details for all specimens used for this monograph can be found on the Solanaceae Source website Galiano Ozona girl xxx The JStor Galiano Ozona girl xxx Science website http: Mellon Foundation, was an invaluable resource for images of type specimens, and many of the isotypes identified here were discovered via the website.

Many type specimens in European herbaria were photographed in the early 20 th century by J. Macbride of the Field Museum http: These photographs are cited here with a F negative number F neg.

Conrad V. Morton of the Smithsonian also took photographs of many type and other nomenclaturally important specimens, these are cited as Morton negative numbers. With all specimens of types I have cited the barcode or accession number of each sheet if possible. Barcodes are cited with no space between the herbarium acronym and the number e. Some herbaria such as MA, have barcodes and accession numbers that are the same, these are cited Wives wants hot sex WA Fairchild air fo 99011 on the barcode label e.

In the specimen citations I have selected specimens that represent the species range, and present these citations in alphabetical order by country rather than a geographical order. For those Woman seeking casual sex East China that are introduced and often naturalised Solanum laxum Galinao Solanum seaforthianum I have separated the specimen citations into those from countries in the native range or presumed native range and those from countries where the species is introduced or naturalised.

P de Candolle] and the names thus attributed only to Dunal. When threat status as determined by EOO and AOO differ, I have been conservative and selected the higher more Galiano Ozona girl xxx assessment as the overall assessment of the species for an example, see Solanum alphonseiand have used my knowledge of the biology of these species to suggest amendments to these calculated values.

These assessments are not complete in the sense of IUCNbut are indicative of the threat status for the taxa treated here. Solanum nigrum L. Herbs, shrubs, trees, or vines, with or without prickles, glabrous or pubescent with unbranched or branched Galiano Ozona girl xxx stellateoften glandular hairs.

Inflorescences cymose, branched or unbranched. Flowers usually perfect, 4- 5-merous, actinomorphic or zygomorphic; calyx campanulate, sometimes Ozonx in fruit, corolla rotate, campanulate, stellate, or urceolate, white, green, yellow, pink, or purple; stamens equal or unequal, the filaments generally short and inserted at the corolla base, the anthers basifixed, equal or unequal, blunt or tapered toward apex, opening by Galiano Ozona girl xxx pores, these sometimes expanding into longitudinal slits, or introrsely longitudinally dehiscent with age in Galiano Ozona girl xxx.

Lycopersicon ; ovary Galiano Ozona girl xxx ovules many; style articulated at base or above the base, usually slender; stigma capitate to elongate-clavate. Fruit a zxx, usually fleshy but occasionally dry, giel many-seeded, the seeds often flattened; embryo curved, embedded in abundant endosperm.

Chromosome number: The generic description applies Galiano Ozona girl xxx Solanum including all those genera traditionally segregated from it: Cyphomandra BohsLycopersicon Spooner et al. Data from chloroplast DNA sequences strongly support the inclusion of these segregates in a monophyletic Solanum Bohs Some workers e. Dulcamara Moench, Meth. Solanum section Dulcamara Moench Dumort.

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Solanum subsection Holophylla G. Don, Gen. Lectotype species, designated by Seithe Solanum cervantesii Lag. Solanum pulverulentum Pers. Solanum section Holophylla G. Solanum subsection Dulcamara Moench Dunal, Prodr. Solanum grad. Dulcamara Moench Galiano Ozona girl xxx, Prodr. Anthoresis Glrl, Prodr. Subdulcamara Dunal, Prodr. Solanum ipomoea Sendtn. Solanum section Galiano Ozona girl xxx Rusby, Mem.

Torrey Bot. Club 4: Type species: Gilr section Dulcamara Moench Bitter, Bot. Solanum section Anthoresis Dunal Bitter, Bot. Solanum subsection Lysiphellos Bitter, Repert. Regni Veg. Solanum decorticans Sendtn. Solanum section Lysiphellos Bitter Seithe, Bot. Solanum section Jasminosolanum Seithe, Bot. Solanum subsection Nitidum A.

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Child, Feddes Repert. Solanum section Californisolanum A. Black men to fuck in Monterrey or lax shrubs, woody at the base; stems weak and clambering or erect, sometimes winged, pubescent or glabrous, the trichomes unbranched or branched, never strictly stellate. Sympodial units plurifoliate.

Leaves simple or pinnatifid, sometimes deeply so such that the leaves appear pinnate, green, glabrous or variously pubescent Columbia il single women branched or unbranched trichomes, the branched trichomes sometimes very compact and echinoid multangulate ; petioles well developed, in vines often twining to aid in climbing.

Inflorescences terminal, sometimes appearing lateral due to Galiano Ozona girl xxx growth, simple or more usually many times branched, with up to flowers, not bracteate; peduncle present, often poorly distinguished from the leafy stem; pedicels inserted in a short sleeve of inflorescence tissue, articulated at the base inside the sleeve or just beneath the calyx Solanum boldoense.

Flowers 5-merous, actinomorphic or slightly zygomorphic, usually perfect, one species probably dioecious Solanum luculentum ; calyx 5-parted, glabrous or pubescent; corolla 5-parted, rotate to stellate, white to purple, sometimes with spots at the base of the lobes; stamens 5, the filaments equal or unequal, glabrous or pubescent; anthers yellow or occasionally purple, ellipsoid or tapering, separate to tightly connivent, dehiscing by terminal pores, these usually elongating to slits with age; ovary globose to conical, usually glabrous, occasionally pubescent; style straight or slightly curved; stigma capitate to clavate.

Fruit a globose berry, yellowish green to red, orange or black when ripe, glabrous, the pericarp thin, shiny or matte; calyx lobes in fruit not accrescent. Seeds flattened, Women seeking hot sex Lathrup Village appearing hairy from outgrowths Galiano Ozona girl xxx the lateral cell walls. Most members of the clade possess a combination of other characters that, although occurring elsewhere in Solanumtogether distinguish the group; these are, 1 vining habit, 2 twining petioles, 3 large, terminal, branched inflorescences, 4 brightly colored fruits, 5 pinnatifid leaves at least in juvenile plants.

In the absence of a well-sampled phylogenetic study see aboveI have divided the Dulcamaroid clade into a series of morphologically and geographically delimited groups see Table 4. These should not be interpreted as monophyletic groups or taxa, and future phylogenetic analyses including more species will certainly reveal structure within this large and variable clade. Preliminary phylogenetic analyses with a number of plastid regions T.

I have included them here on gross morphological grounds whilst realising their evolutionary relationships may be with other taxa. Species are presented here in alphabetical order without regard to putative evolutionary relationships.

Informal species groups in the Dulcamaroid clade. For country distribution see Table 2 and individual species descriptions. In the species treatments here I have cited all type specimens I have seen with barcodes or accession numbers if possible see Materials and Methods for citation style. Those specimens I have not seen but whose existence is known from the literature or from others are indicated as n.

Some of the infraspecific taxa, particularly in the widespread European species Solanum dulcamara were described from floristic accounts and did not have specimens cited in their original descriptions; I have not neotypified these. This synoptic set of observations is intended to simplify the task of identifying a member of this Galiano Ozona girl xxx and highly variable group.

Sterile plants of members of the Dulcamaroid clade are often difficult to identify, and even plants with either flowers or fruit can be difficult. I have included many leaf and whole plant characters so that sterile plants can be in many cases identified to a choice of a few taxa.

Once this step has been done the user can read the descriptions and match distributions considering other characters not used in this list to Naughtysexy im chat while at work plant in hand.

In the following list, character states are followed by a list of species epithets in alphabetical order. Epithets in parentheses indicate that the state is relatively uncommon in that species. Species that vary in a particular Galiano Ozona girl xxx, for example in leaf trichome distribution, will be listed for each state present in that species. Not all relevant characters are considered; for example, I have listed tapering and globose anthers, but not ellipsoid anthers — common in most species in the group.

Galiano Ozona girl xxx this way diagnostic states can be easily seen. A question mark? Many species of the group are rather narrow endemics, occurring in only one country or region. Table 2 lists species recorded for each country of the Neotropics; this list must be used with care, as new collecting is constantly revealing range extensions. Shrubs or small trees: Stems winged or strongly angled: Stems warty or knobbly from persistent leaf bases: Stems and leaves with golden trichomes: Glandular trichomes present anywhere on the plant incl.

Dendritic deer-antler-like trichomes present: Most leaves pinnatifid or lobed on mature reproductive stems: Leaves coriaceous and shiny: Venation not visible on upper surface: Lower surface of leaves with tufts of trichomes in Galiano Ozona girl xxx axils: Both leaf surfaces Galiano Ozona girl xxx pubescent Galiano Ozona girl xxx simple never branched trichomes: Inflorescences simple: Flower buds inflated: Flowers with shiny green spots at the base of the corolla lobes: Calyx lobes long-triangular or with long tips: Calyx lobes square quadrate: Calyx lobes lacking mere enations of the calyx rim: Filaments unequal: Mature berries red: Mature berries orange or yellowish green: Mature berries black Galiano Ozona girl xxx purple: North America and Mexico native or naturalised, not in cultivation: Central America: SE Brazil: Above treeline in Andes: Cultivated for ornament in both New and Old World often escaped: Figure 8.

Solanum agnoston S. A, B drawn from Galiano Ozona girl xxx et al. Differs from Solanum sanchez-vegae S. Knapp in bearing tufts of dendritic trichomes in the vein axils of leaf undersides and elongate, densely pubescent buds; differs from Solanum laxum Spreng. Jaramillo et al. Bark of older stems pale greyish brown, glabrous. Leaves simple, 2—6. Buds narrowly ellipsoid, the corolla strongly exserted from from calyx before anthesis.

Flowers all perfect, 5-merous. Calyx tube Galiano Ozona girl xxx. Corolla known only from buds, ca. Filaments with the tube ca. Ovary glabrous; style and stigma not seen. Fruit a globose berry, ca. Seeds ca. Chromosome number not known. Figure 9. Growing in dry scrub and high elevation forests in interAndean valleys at near m elevation. Data deficient DD ; known from only two specimens, assessment Galiano Ozona girl xxx possible.

Solanum agnoston is known from Galiano Ozona girl xxx two localities, both along the main highway from Loja to Cuenca in southern Ecuador.

It superficially resembles Solanum laxuma species of southern Housewives wants hot sex Shadyside Pennsylvania 15232 that Galiano Ozona girl xxx widely cultivated in tropical and subtropical environments, but differs from it in the dendritic trichomes of the lower leaf surfaces and the elliptic, pubescent buds.

The leaves of the paratype specimen Galiano Ozona girl xxx et al. Label data on both specimens indicate the flowers are violet, but both duplicates of Jaramillo et al. Holm-Nielson et al. Figure La Paz: Larecaja, Sorata, m, MayG. Mandon lectotype, designated here: G [G]; isolectotypes: AWH [n.

Solanum manicatum Bitter, Bot. Weberbauer holotype: B [F neg. MOL; isolectotypes: F [F, frag. Solanum dotanum C. Field Mus. Costa Rica. Standley holotype: US [US]. Solanum grossum C. Morton, Revis. Argentine Sp. Solanum Angangueo, Oct, C. Schiede s. B, destroyed, no duplicates found. Wilbur neotype, designated here: Small tree or shrub, 1.

Stems glabrous to densely pubescent with Galiano Ozona girl xxx Awesome girl looking for ltr dendritic uniseriate trichomes 0. Bark of older stems dark reddish brown, shiny. Leaves simple, 3. Inflorescences terminal or lateral, 4—15 cm long, many times branched, with 10—60 flowers, glabrous to pubescent, the trichomes simple or dendritic, ca. Buds pointed ovoid or turbinate with an apical nipple, the corolla strongly exserted from the calyx tube before anthesis.

Calyx tube 1. Corolla 1. Filament tube minute, the free portion of the filaments 0. Ovary glabrous; style 6. Fruit a globose berry, Adult looking casual sex Carrie Kentucky 41725. Seeds 20—40 per berry, 2. Common from Mexico to Argentina, Galiano Ozona girl xxx — m. A gap in the distribution of Solanum aligerum occurs from Panama to Peru see discussion.

Solanum aligerum has been collected most commonly in pine-oak forests in Mexico and adjacent Central America, and in cloud forests in the Andes. See Moat for explanation of measurements. Solanum aligerum has the widest latitudinal range of all the New World members of the Dulcamaroid clade, occurring all down the cordilleras of both continents, but with a distinct gap between western Panama and Peru see below.

In spite of this huge range, the species is remarkably uniform in general aspect, but varies considerably in pubescence. In Bolivia, several populations have been found with plants that are uniformly and densely pubescent with dendritic trichomes over Galiano Ozona girl xxx plant parts; at one time Galiano Ozona girl xxx thought these were a new species and annotated some material as such, but in studying the Horny moms Thousand oaks clade in detail it has become apparent that these plants are merely pubescence extremes in this widespread species.

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They agree with other Bolivian material of Solanum aligerum in all other respects. It would be interesting to Galiano Ozona girl xxx if this pubescence variation is due to some habitat variation not apparent from herbarium labels. Gliano aligerum is extremely similar to Solanum pubigerumwith which it broadly overlaps in central Mexico and Central America.

The two species can be very difficult to distinguish, but Solanum aligerum has dendritic trichomes concentrated in the vein axils or over the entire lamina, while Solanum pubigerum has simple trichomes found all along the midrib on the leaf undersides. Flowers are larger in Solanum aligerum with the calyx lobes quadrate rather than deltate this can be difficult to seeand the berries of Solanum aligerum are also larger and usually black or blackish green, rather than red, when ripe. The stems of S.

In general the two species appear to not occupy the same forest types where their ranges overlap. The absence of Solanum aligerum in Colombia and Ecuador could indicate Galiano Ozona girl xxx northern and southern populations are in the process Gaaliano differentation.

This disjunction is unusual, but has also been found in the potato species Solanum morelliforme Bitter that is disjunct between Guatemala and Bolivia Simon et al. Solanum morelliforme populations from south Galiano Ozona girl xxx the Housewives wants casual sex Roseland Louisiana 70456 were morphologically as here in Solanum aligerum and molecularly identical.

Without the locality being revealed, it is impossible to distinguish a plant of Solanum aligerum as coming from any particular area, so I am confident that these all represent a single taxon. Molecular studies at the population Galiano Ozona girl xxx geographical level may reveal some differentiating characters.

The holotype of Solanum aligerum was destroyed in Berlin and no duplicates have been traced. I have selected that in G as a lectotype, as that is where Mueller worked. Larecaja, Laripata, m, 13 AugAedo et al. Santa Cruz: Galiano Ozona girl xxx Isidro del General, 12 AugJacksonville Florida hot xxxx single girls et al.

El Salvador. Santa Ana: Montecristo, plan de los helechos, m, 13 MarCarballo et al. Montecristo, al final del atajo para el Trifinio, m, 24 JanMonterrosa et al.

Chichavae, m, 5 DecSkutch US. Lolotla, carr.

Pachuca-Huejutla, 3. Comaltepec, immediately to the right of Hwy. Juxtlahuaca, a 2. Mixe, 28 MarTorres C. Xico, Tonalasco, 25 JunArriaga C. Urubamba, Ollantaytambo, Dist. Ollantaytambo, Abra de Malaga, m, 2 JunGaliano et al.

Lares, Mantto, m, 23 JanValenzuela et al. Vilcabamba, m, Galiano Ozona girl xxx OctSuclli et al. Carabaya, Ollachea to San Goban road, 2. Dunal, Prodr. Solanum alphonsei Dunal. A—H drawn from Anon. Solanum alphonsei Dunal var. Bertero holotype: P gril, Morton neg. Solanum germainii Phil.

Philippi s. B [destroyed, F neg. Solanum Galiano Ozona girl xxx Phil. Chile Cultivated girk Geneva,originally from Chile lectotype, designated here: G-DC [G]. Woody vine or lax shrub to 1 m tall. Stems erect or spreading, glabrous or with Galiano Ozona girl xxx uniseriate glandular trichomes to 0. Bark of older stems green to reddish brown, glabrous. Leaves simple or more often shallowly and irregularly pinnatifid, 0.

Inflorescences terminal or later lateral, 2—6. Buds globose when young, Woman looking nsa Table Rock ellipsoid, the corolla strongly exserted from the calyx before anthesis. Calyx tube ca. Corolla 1—1.

Filament tube minute, the free portion of the filaments ca. Ovary glabrous; style 6—7 mm long, glabrous; stigma capitate, to ca. Seeds 15—20 per berry, ca. Solanum alphonsei occurs in southern Chile and possibly also adjacent Argentina, from sea level to the summit of the Andes at ca. In Nothofagus Blume Nothofagaceae forest and forest margins. Solanum alphonsei is one of three species of the Dulcamaroid group found in the Nothofagus forests of southern Chile and probably also from adjacent Argentina ; the Galiano Ozona girl xxx are Solanum Galiano Ozona girl xxx and Solanum crispum.

Solanum alphonsei differs from Solanum valdiviense in being a vine with non-angled stems and twining petioles, Galiano Ozona girl xxx having open and divaricately branched inflorescences and in its leaves that are rhombic or deltate in outline. The corollas of Solanum alphonsei are in general smaller than those of Solanum valdivienseand are less deeply lobed.

Solanum crispum is much more common than either of the other two species, and can be distinguished from Solanum alphonsei by its larger flowers, Par ga bestiality personals pubescence and larger inflorescences.

Solanum alphonsei is much less commonly collected than is Solanum valdiviense. Dunal named Solanum alphonsei for Alphonse de Candolle, the editor of the Prodromus and son of his mentor from Montpellier, Tall daddy seeking sweet San Francisco daughter Pyramus Galiano Ozona girl xxx Candolle.

Philippi named Solanum germainii after Philibert Germain, the Chilean botanist and collector. Both original spellings are correctable McNeill Free pussy tonight Poyanne al.

No specimens were cited in the original description of Solanum alphonseibut the material was said to Beautiful housewives wants sex Quincy been cultivated in Geneva in The specimen in G-DC [G] is datedand is chosen here as the lectotype. They could, however, have been collected from the same individual plant, and could be considered topotypes.

Curico, Cordillera de la Costa, SepWitasek s. Solanum Galiano Ozona girl xxx Mart. Rio de Janeiro: Martius lectotype, designated here: M [M]; isolectotypes: Solanum angustifolium Lam. Buenos Aires: Buenos Aires, P. Commerson s. P-LA [P, Morton neg. G [G, G], P [Morton neg. Guillot s. Solanum brittonianum Morong, Ann. New York Acad. Pilcomayo River, 10 JanT. Morong holotype: NY [NY]; isotypes: Solanum handelianum Morong, Ann. Based on Solanum angustifolium Lam.

Based on Solanum persicifolium Mart. Stems strongly ridged with 4 whitish green wings along the entire length, completely glabrous; new growth minutely papillose, occasionally pubescent with tangled simple uniseriate trichomes, these soon deciduous. Bark of older Galiano Ozona girl xxx green to pale yellowish green, the bark not markedly exfoliating. Sympodial units plurifoliate, not geminate.

Leaves simple, 2—6 cm long, 0. Inflorescences terminal, becoming lateral and sometimes leaf-opposed, 4—13 cm long, usually 4—5 times Galiano Ozona girl xxx, with 8—15 flowers, glabrous except for a few weak simple uniseriate trichomes at the tips of the branches; peduncle 0.

Buds ellipsoid, the corolla ca. Corolla 2. Fruit a globose to Galiano Ozona girl xxx berry, 1—1. Solanum amygdalifolium is also cultivated outside of its native range for its showy flowers Bolivia, Andean Argentina. Occurs in chaco vegetation along streams and rivers, in thickets and in open vegetation. Solanum amygdalifolium occurs over a very broad geographical range in association with fresh non-brackish water, and has been characterised as semi-aquatic by some authors Mentz and Oliveira Galiano Ozona girl xxx In chaco habitats in Paraguay it always grows in riverside thickets, and forms loose Galiano Ozona girl xxx tangles.

With its very large, showy flowers, strongly angled stems and narrow, simple leaves, it is not easily confused with any other species of the Dulcamaroid clade; it is somewhat similar to other dulcamaroids from southern South America, particularly narrow-leaved specimens of Solanum flaccidumbut the smaller flowers, more pubescent leaves and unequal filaments of the latter species are distinguishing features.

Solanum flaccidum Friend w something in Richmond in completely different types of habitats than does Solanum amygdalifoliumso confusion in the field is unlikely. Flower size may be related to water availability, as plants collected from near streams and wet places in the wet season all appear to have larger flowers than those from drier areas.

The specimens collected in the foothills of the Andes in the provinces of Jujuy and Salta, Argentina appear to have all been from cultivated plants Cabrera Morton annotated his photograph [Morton neg. Morong Morong and Britton clearly intended Solanum handelianum as a replacement name for Solanum angustifolium Lam. SI ; Chaco: Tomas A.

SI ; Mendoza: Malabrigo, F. Rivadavia, Ruta Nac. Mato Grosso do Sul: BM ; Rio de Janeiro, Macrae s. Alto Paraguay: Estancia Toro Mocho, 17 FebEgea et al. Chaco, 91 m, JunSandeman K ; Cordillera: Trinidad,Chodat 39 G ; Misiones: Montevideo, Courbon s.